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Ryan Harnish

Pacific Northwest National Laboratory
PO Box 999
Richland, WA 99352


Ryan Harnish is a research scientist. Since joining PNNL in 2007, his work has primarily focused on the use of acoustic telemetry to assess the behavior and survival of salmon and steelhead smolts in the Snake and Columbia River basins. Specifically, this work has addressed the effects of hydropower projects on the behavior and survival of smolts, and the effects of acoustic transmitter implantation on the performance of tagged fish.

Research Interests

  • Acoustic telemetry
  • Survival and behavior of salmonid smolts
  • Effects of hydropower projects on fisheries resources
  • Effects of irrigation on fisheries resources
  • Conservation of native fishes

Education and Credentials

  • M.S., Fish and Wildlife Management, Montana State University, 2007
  • B.S., Biology, Montana State University, 2001

Affiliations and Professional Service

  • American Fisheries Society

PNNL Publications


  • Deng Z, JJ Martinez, H Li, RA Harnish, CM Woodley, JS Hughes, X Li, T Fu, J Lu, GA Mcmichael, MA Weiland, MB Eppard, JR Skalski, and RL Townsend. 2017. "Comparing the survival rate of juvenile Chinook salmon migrating through hydropower systems using injectable and surgical acoustic transmitters." Scientific Reports 7:42999.  doi:10.1038/srep42999
  • Mueller RP, JM Janak, SA Liss, RS Brown, Z Deng, and RA Harnish. 2017. "Retention and Effects of Miniature Transmitters in Juvenile American Eels." Fisheries Research 195:52-58.  doi:10.1016/j.fishres.2017.06.017




  • McMichael GA, AC O'Toole, RA Harnish, and DM Trott. 2013. "Juvenile salmonid migratory behavior at the mouth of the Columbia River and within the plume." Animal Biotelemetry 1:Article No. 14.  doi:10.1186/2050-3385-1-14


  • Harnish RA, GE Johnson, GA McMichael, MS Hughes, and BD Ebberts. 2012. "Effect of Migration Pathway on Travel Time and Survival of Acoustic-Tagged Juvenile Salmonids in the Columbia River Estuary." Transactions of the American Fisheries Society 141(2):507-519.  doi:10.1080/00028487.2012.670576
  • Mueller RP, EV Arntzen, MA Nabelek, BL Miller, KJ Klett, and RA Harnish. 2012. "Laboratory testing of a modified electroshocking system designed for deepwater juvenile lamprey sampling ." Transactions of the American Fisheries Society 141(3):841-845.  doi:10.1080/00028487.2012.675908


  • McMichael GA, RA Harnish, JR Skalski, KA Deters, KD Ham, RL Townsend, PS Titzler, MS Hughes, JA Kim, and DM Trott. 2011. Migratory Behavior and Survival of Juvenile Salmonids in the Lower Columbia River, Estuary, and Plume in 2010 PNNL-20443, Pacific Northwest National Laboratory, Richland, WA. 
  • McMichael GA, RA Harnish, MA Weiland, Z Deng, and MB Eppard. 2011. "Fish Passage: A New Tool to Investigate Fish Movement: JSATS." Hydro Review 30:34-42. 
  • Harnish RA, AHA Colotelo, and RS Brown. 2011. "A review of polymer-based water conditioners for reduction of handling-related injury." Reviews in Fish Biology and Fisheries 21(1):43-49.  doi:10.1007/s11160-010-9187-1


  • McMichael GA, MB Eppard, TJ Carlson, JA Carter, BD Ebberts, RS Brown, MA Weiland, GR Ploskey, RA Harnish, and Z Deng. 2010. "The Juvenile Salmon Acoustic Telemetry System; A New Tool." Fisheries 35(1):9-22. 
  • McMichael GA, RA Harnish, BJ Bellgraph, JA Carter, KD Ham, PS Titzler, and MS Hughes. 2010. Migratory Behavior and Survival of Juvenile Salmonids in the Lower Columbia River and Estuary in 2009 PNNL-19545, Pacific Northwest National Laboratory, Richland, WA. 
  • Brown RS, RA Harnish, KM Carter, JW Boyd, KA Deters, and MB Eppard. 2010. "An evaluation of the maximum tag burden for implantation of acoustic transmitters in juvenile Chinook salmon." North American Journal of Fisheries Management 30:499-505. 




Additional Information